Template:Dwyer, 1980
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- Dwyer, J.D. 1980. Family 179. Rubiaceae—Part 1. In Woodson, R.E., Jr., Schery, R.W. & Collaborators, Flora of Panama, Part IX. Annals of the Missouri Botanical Garden 67: 1–256. DOI: 10.2307/2398954 BHL JSTOR